They are the exact same car. The EPA fuel economy testing procedure changed for the 2008 model year. ALL cars are measured in a laboratory under a specified driving pattern by the EPA. The new 2008 testing procedures adds in extra testing for higher speeds/acceleration, colder temperatures, and AC use, which the older tests did not. ALL cars, not just the Prius, had their MPG go down in 2008.
http://www.fueleconomy.gov/feg/ratings2008.shtml
http://www.fueleconomy.gov/feg/fe_test_schedules.shtml
http://www.fueleconomy.gov/feg/how_tested.shtml
US 2004-2007 EPA ratings were: 60MPG city, 51MPG highway, 55MPG combined.
Under the new EPA testing schedule, the 2008 (and the 2004-2007 have been revised to be): 48MPG city, 45MPG highway, 46MPG combined.
Toyota, like all other manufacturers, are REQUIRED by law to publish the EPA test numbers on the window stickers. So don't blame Toyota, blame the EPA for having such outdated and unrealistic tests. (The EPA just added some new tests for the 2008 model year to help with the MPG inflation that ALL cars, not just the Prius/hybrids, used to have. All cars' reported 2008 MPG have dropped, by about 15-20%.) The EPA tests are done the same to all cars, so that you can compare one car against another under the same driving cycle. Governmental test cycles are really only good for comparing one car against another (standardized tests), and not what you can expect in your real-world driving (as your idea of "city" driving may not be what the government test uses as "city" driving).
Your average fuel economy is same/better than what other owners report:
https://www.fueleconomy.gov/mpg/MPG....ta&model=Prius and
http://www.greenhybrid.com/compare/m...-priushsd.html