The Lock Button Turns on 4WD, in normal driving conditions, the 4WD will turn on and off when needed. "Trac" means traction control, bascially it's what you turn ON when you're just driving around in case of an emergency (Unless you're Lewis Hamilton, but in that case, Why would you be driving a RAV4?). When going offroading it helps to turn traction control off because it gives you more control at low speeds.
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